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Primary forests are frequently perceived as undisturbed forest
in a late stage of succession, but the concept should include
the associated younger successional stages. In contrast, secondary
forests are defined as forests regenerating largely through
natural processes after significant human disturbance of the
original forest at a single point in time or over an extended
period and displaying a major difference in forest structure
and/or canopy species composition with respect to nearby primary
forests on similar sites. Because of the large scale and intensity
of the disturbance, secondary forests pass through stages
in a succession series that is different from the typical
forest vegetation of that site.
While conservationists traditionally focus on primary forests
for their biodiversity and natural processes, there is much
recent attention for secondary forests, not in the last place
because primary forests are more and more scarce. This comes
with the appearance of large and growing areas of secondary
forests in areas formerly used for agriculture or intensive
logging. Recent estimates indicate the existence of 500-600
million ha of secondary forests in the tropics. It is widely
expected that future goods and services from tropical forests
will increasingly be met by secondary forests (in addition
to plantations), including biodiversity conservation. There
are several types of secondary forest. Among the most significant
are the areas created by commercial logging and secondary
forests that grow on land that has been used by shifting cultivators.
Shifting cultivators often create secondary forest gardens
by planting fallow areas with trees that provide fruits, nuts,
resins and other products. Fire leads to yet another type
of secondary forest succession.
Being the product of a heavy disturbance, secondary forests
undergo a succession along the lines described in the section
on forest dynamics. In the literature,
three general phases of succession in secondary forests are
distinguished. In the first phase, colonisation occurs by
a dense growth of herbs, shrubs and climbers. Within 1-2 years,
these will be replaced by short-lived pioneers: trees with
a life span of 10-30 years. These form an even canopy, under
which seedlings of long-lived pioneers and climax species
slowly grow. After the death of the short-lived pioneers,
these will form a canopy which is still relatively uniform
and even-aged, and which may transform into a forest resembling
mature forest after possibly 75 to 150 years. During this
succession, species richness may build up quite rapidly and
reach levels comparable to mature forests in 20 to 80 years.
However, species composition is quite different from mature
forests, mainly through the abundance of short- and long-lived
pioneers, which are naturally scarce in mature forest.
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TBI sources
Vester,
H.F.M. (1995). The trees of the forest. The role of tree
architecture in canopy development; a case study in secondary
forest (Araracuara, Colombia).
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